For his PhD, John studied in Eric Schmidt’s laboratory at the University of Utah, where he studied the biosynthesis of the cyanobactin family of cyclic peptide natural products.

After finishing graduate school John was a postdoc in Frances Arnold’s laboratory at Caltech where he studied non-natural reactions of cytochrome P450s and other heme proteins.

John have been with Merck’s Small-Molecule Process Research and Development Department since 2015 where he has worked in the Biocatalysis and Protein Engineering groups. He is presently an associate principal scientist.